Thanks, Michael, and good morning to everyone. Michael has already alluded to our financial performance in 2022. And now I'd like to talk further about our first quarter 2020 results. Net income of $3.1 million in the first quarter of 2022 was lower by $2.2 million, mainly due to a decline in gains on sales of loans, lower net interest income and lower gains on sales of investment securities.
In the first quarter of 2022, net interest income totaled $8.6 million, a decrease of $946,000 or 9.9% in comparison to the same period last year. While on a linked quarter basis, net interest income was down by $491,000. The decline in net interest income from the first quarter last year as well as the fourth quarter of 2021 was mainly the result of decreased interest this quarter on Paycheck Protection Program, or PPP and other commercial-related loans, but partly offset by lower deposit costs.
Interest on PPP loans in the first quarter of 2022 declined by $636,000 compared to the first quarter last year, while the PPP balance interest on the PPP balance loans declined by $112.1 million over the same period. The average tax equivalent yield on the loan portfolio has declined this quarter to 4.59% compared to 4.67% in the same period last year and 4.81% last quarter.
Interest income on investment securities increased to $186,000 this quarter compared to the same period last year due to a growth in average investment balances of $120.4 million, but offset by lower yields. The yield on investment securities declined from 2.37% in the first quarter 2021 to 1.83% in the current quarter, which was an increase from 1.77% in the fourth quarter of 2021.
The investment portfolio growth in the first quarter of 2022 resulted from deploying excess cash balances into investments with a focus on shorter duration U.S. treasuries that should perform well in a rising rate environment. Interest costs on interest-bearing deposits remained low this quarter, totaling 10 basis points in the current quarter compared to 15 basis points in the first quarter of 2021 and 12 basis points last quarter. Interest expense on total deposits declined $86,000 from the first quarter of last year due to lower rates, offset by growth in average balances of $29.6 million in interest-bearing deposits.
Landmark's net interest margin on a tax equivalent basis decreased to 2.99% in the first quarter of 2022 as compared to 3.17% in the fourth quarter of 2021. Our loan-to-deposit ratio, which totaled 55% at March 31, 2022, remains low, giving us plenty of opportunities to fund new loan growth.
Based on our analysis of the economic environment, our strong credit results and a decline in loans this quarter, excluding PPP loans, we recorded a $500,000 reverse provision to the allowance for loan losses in the first quarter of 2022. At March 31, 2022, the ratio of our loan loss reserve to gross loans, excluding PPP loans at year-end was 1.33%. As our economic outlook evolves, we will continue to adjust our allowance for credit losses and provisioning accordingly.
Noninterest income totaled $3.6 million this quarter, decreasing $3.2 million compared to the first quarter of 2021, while declining by $1.0 million in comparison to the prior linked quarter. This decrease over the same period last year was due mainly to a decline of $2.2 million in sales of 1 to 4 family real estate loans that the bank originated.
During the current quarter, higher interest rates, coupled with a lack of housing inventory in our markets, slowed purchase and refinancing activities as compared to the first quarter last year when mortgage activity was extremely strong.
The first quarter of 2021 included a gain of $1.1 million on the sale of higher coupon municipal investment securities that did not occur in the current quarter. These declines were offset by an increase over the same quarter last year of $155,000 in fees and service charge income.
Noninterest expense for the first quarter of 2022 totaled $8.8 million, representing a decrease of $235,000 over the same period last year and was $712,000 lower than the prior quarter. The decrease over the first quarter of 2021 was driven by a decline of $166,000 in compensation and benefits, primarily related to lower mortgage lending activities, along with declines of $161,000 in data processing and $121,000 in amortization expense. These decreases in noninterest expense were partially offset by an increase of $171,000 in occupancy and equipment.
The effective tax rate was 19.0% in the current quarter, down from 20.4% in the first quarter of 2021. I Total assets remained stable during the first quarter at $1.3 billion for each of the 2 recent quarter ends. Gross loans, excluding PPP loans, decreased $16.9 million during the first quarter, and as mentioned, related to reductions in both agricultural lending and other commercial loans.
Our deposits decreased by $8.9 million during the quarter to $1.1 billion due in part to seasonality of public funds, while the decrease in cash and cash equivalents of $82.9 million funded growth in investment securities of $86.3 million.
Stockholders' equity decreased to $123.5 million at March 31, 2022, and our book value decreased to $24.72 per share. The decrease in book value was due to a decline in the fair value of our investment securities, which were impacted by higher interest rates.
Our consolidated and bank capital ratios as of March 31, 2022, are very strong and exceed the regulatory levels considered to be well capitalized. The bank's leverage ratio was 10.5% at March 31, 2022, while the total risk-based capital ratio was 19.2%.
Now let me turn the call over to Raymond to review highlights of our loan portfolio and the credit risk outlook.